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Cluster info 4grp_rule_based, cluster L2

   > (GO:0005575) cellular_component [514, 0] [dist]
         > (GO:0031974) membrane-enclosed lumen [8, 0] [dist]
               -> (GO:0031970) organelle envelope lumen [1, 0] [dist]
               -> (GO:0043233) organelle lumen [8, 0] [dist]